Xinyu Liu1,2,3, Selin Scherrer4, Sven Egger4, Song-I Lim2, Benedikt Lauber4, Wolfgang Taube4, and Lijing Xin2,3
1Laboratory for functional and metabolic imaging (LIFMET), Ecole Polytechnique Fédérale de Lausanne, Lausanne, Switzerland, 2Center for Biomedical Imaging (CIBM), Lausanne, Switzerland, 3Animal Imaging and Technology, Ecole Polytechnique Fédérale de Lausanne, Lausanne, Switzerland, 4Department of Neurosciences and Movement Science, University of Fribourg, Fribourg, Switzerland
Synopsis
Keywords: Aging, Nervous system, Cortical inhibition
Motivation: The effect of short-term physical activity on brain metabolism has been widely studied in young population, but less is known about metabolic plasticity induced by strength training over a long range among elderly adults.
Goal(s): This study examined the modulatory potential of long-term strength training on metabolism in elderly population.
Approach: Using 7 tesla magnetic resonance spectroscopy (MRS), we assessed concentrations of γ-Aminobutyric acid (GABA), glutamate (Glu), and a number of other metabolites in the sensorimotor cortex.
Results: We found that three-months of strength learning significantly reduced lactate levels in the sensorimotor cortex compared to a passive control group.
Impact: This study systematically examined brain metabolic
plasticity induced by strength training in the elderly population. The
reduction in lactate indicates that strength training may facilitate oxidation
of lactate to meet increased energy demands for learning.
Intoduction
Physical exercise is an important measure
to counteract age-associated physiological deterioration processes. Acute
physical activity has been shown to modulate metabolism in the human brain, such
as elevation of GABA, Glu1 and
lactate2 in
visual cortex. However, most of previous studies focused on young population
and it remains elusive whether similar modulatory effects can be observed after
a long period of physical training, especially in elderly population who are
known to suffer from reduced physical abilities.
In
this study, we seek to evaluate the effects of long-term strength training on brain
metabolism in elderly adults using 7T magnetic resonance spectroscopy (MRS). The
concentration of the main inhibitory neurotransmitter GABA were measured by
MEGA-sSPECIAL, a method reported to reliably quantify metabolites GABA levels
in sensorimotor cortex with high reproducibility3. Semi-adiabatic
SPECIAL (sSPECIAL) sequence was used for neurochemical profiling of other
metabolites.
Methods
Thirty-two healthy volunteers (66 – 82
years old, 17 males/15 females) gave informed consent prior to the study. They
are randomly assigned to strength learning (n = 16) or a control group (n =16).
We used MP2RAGE T1 weighted images for
voxel positioning, MEGA-sSPECIAL for edited GABA measurement, and sSPECIAL for
neurochemical profiling. Detailed acquisition parameters can be found in Table
1. After the first MRS session, participants in the strength training group
underwent three months of progressive, multifaceted strength training. They were
trained around 3 times per week (a total of at least 30 training sessions) for
45 minutes in supervised group sessions. Participants in the control group did
not engage in any training. After three months, both groups underwent the
second measurement session with identical MR protocol as the first one.
MR spectra were averaged after frequency
drift and phase correction using FID-A4 and
analyzed by LCModel5 for
quantification. Metabolites with Cramer-Rao lower bound (CRLB) lower than 30%
were reported. Unsuppressed water signal was used for quantification and water
content was corrected for tissue composition. Metabolite concentrations were
corrected for CSF contribution. A linear mixed-effect model with group as
between-subject factor and time as within-subject factor was used to detect
whether there is an interaction effect between time and group. False discovery
rate (FDR) correction were performed using Benjamini and Hochberg procedures6
when comparing metabolite concentrations detected using sSPECIAL. When there is
a significant interaction effect, post-hoc two-sample t-tests were followed.Results
Demographic information and spectral
quality parameters can be found in Table 2. There is no significant
difference observed in signal-to-noise ratio (SNR) and linewidth between the
pre- and post-measurements.
Figure 1 showed an exemplar voxel placement and spectra for left
sensorimotor cortex.
Using linear
mixed-effect model, we found that there is no significant interaction effect of
time × group in GABA level (p = 0.14, z59 = -1.493, [-0.730, 0.099])
(Figure 2A). Semi-adiabatic SPECIAL detected eleven metabolites of interest. We found
that there is a significant interaction effect of time × group in lactate level
(corrected p = 0.049, z57 = -2.839, [-0.762, -0.140]). Post-hoc
analysis suggests that this is driven by a significant decrease in lactate in
strength training group (p = 0.02) but not in the control group (p = 0.33) (Figure 2B). No significant interaction effects were found for other investigated
metabolite concentrations (Figure 3).Discussion
Lactate is an
important energy metabolite in the brain which ensures adequate energy supply,
modulates neuronal excitability levels and regulates adaptive functions in
order to set the 'homeostatic tone' of the nervous system7. Multiple MRS studies have reported increased brain lactate
level following acute physical exercise2,8. Contrast to short-term studies, we found that long-term
strength learning significantly reduced sensorimotor cortical lactate level in the
learning group compared to the control group. Previous animal studies have
shown that lactate accumulation is a hallmark of aging and is associated with
elevated lactate dehydrogenase (LDH)-A/LDH-B ratio9. Thus, one potential explanation for the observed
lactate reduction could be that strength learning can facilitate the oxidation
of lactate in the brain into pyruvate which enters in oxidative phosphorylation
for energy production in aerobic fashion.Conclusion
In conclusion, the
present study found that long-term strength training significantly reduced
lactate level compared to a control group. This may indicate that strength training
has the potential to facilitate oxidation of
lactate in the brain to meet increased energy demand for learning, and shed new
light into the distinct modulatory mechanisms between short-term and long-term
physical exercise on brain metabolism.Acknowledgements
This
work was supported by the Swiss National Science Foundation (grants
n° 32003B_197687). We acknowledge access to the facilities and expertise
of the CIBM Center for Biomedical Imaging, a Swiss research center of
excellence funded and supported by Lausanne University Hospital (CHUV),
University of Lausanne (UNIL), Ecole Polytechnique Fédérale de Lausanne (EPFL),
University of Geneva (UNIGE) and Geneva University Hospitals (HUG).References
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