Radka Klepochova1,2, Fabian Niess2, Siegfried Trattnig2,3,4,5, Alexandra Kautzky-Willer1, Martin Krššák1,2, and Martin Meyerspeer6
1Division of Endocrinology and Metabolism, Department of Internal Medicine III, Medical University of Vienna, Vienna, Austria, 2High-Field MR Center, Department of Biomedical Imaging and Image-Guided Therapy, Medical University of Vienna, Vienna, Austria, 3CD Laboratory for MR Imaging Biomarkers (BIOMAK), Vienna, Austria, 4Austrian Cluster for Tissue Regeneration, Ludwig Boltzmann Institute for Experimental and Clinical Traumatology, Vienna, Austria, 5Institute for Clinical Molecular MRI in the Musculoskeletal System, Karl Landsteiner Society, Vienna, Austria, 6High-Field MR Center, Center for Medical Physics and Biomedical Engineering, Medical University of Vienna, Vienna, Austria
Synopsis
Keywords: Muscle, Spectroscopy
Motivation: To explore quantification of skeletal muscle oxidative metabolism by 1H MRS.
Goal(s): We used the increased accuracy of 7T MRS with a dedicated RF-coil, interleaved acquisition and localization of 31P information to compare Creatine-CH2 and Phosphocreatine time courses during exercise and recovery.
Approach: Eight volunteers were measured on a 7T MR system with RF-coil and ergometer dedicated for exercise. 1H and 31P MR spectra were acquired interleaved during exercise and recovery.
Results: Exercise led to disappearance of the Creatine-CH2 resonance, while the CH3 resonance remained stable during exercise. The recovery time constants were similar (τPCr=37±9s and τCr=34±6s) and positively correlated.
Impact: The
time course of the Creatine-CH2 resonance in skeletal muscle can be accessed
via dynamic 1H MRS. If accurately reflecting oxidative metabolism,
this technique has the potential to render non-invasive metabolic studies
broadly accessible, without needing multi-nuclear MRI capabilities.
Introduction
Localized
1H and 31P MR spectra of human muscle are strongly affected
by exercise. Phosphocreatine (PCr) levels measured by 31P MRS
decrease during exercise, reach steady state, and recover towards basal values
after exercise. 1H MRS exhibits two Creatine (Cr) signals, CH3
(3.02 ppm) and CH2 (3.92 ppm). Exercise-induced PCr depletion is
known to influence also the spectral appearance of the Cr-CH2 3.92
ppm resonance1.
We
used the increased accuracy of 7T MRS with a dedicated RF-coil, interleaved
acquisition and localization of 31P information to compare Cr-CH2
and PCr time courses during exercise and recovery. Furthermore, we investigated
skeletal muscle oxidative metabolite kinetics from time-resolved 1H
MRS.Methods
Twelve
healthy, recreationally active volunteers (age: 30±7 years, BMI: 25.2±7.9kg/m2,
sex: 5f/7m) participated in the study on a 7T whole-body MR system
(Terra-DotPlus, Siemens Healthineers, Erlangen, Germany). A dedicated 1H(2-channel)/31P(3-channel)
surface coil transceiver array2 was used on
the subject’s right calf. An MR-compatible ergometer (Trispect, Ergospect,
Innsbruck, Austria) was used for plantar flexion exercise. Data from four
subjects were excluded in the analysis due to either poor shim (> 70 Hz,
n=2), water-suppression (n=1) or baseline distortion (n=1), all presumably
caused by motion.
Measurements
were performed in the morning after overnight-fasting in a single
exercise-recovery session. Volunteers were positioned supine with the right
calf on the coil and ergometer, inside the scanner.
The
VOI for 1H MRS Cr detection (15x40x53 mm3) and 18-mm slab for 31P MRS PCr and inorganic
phosphate (Pi) detection were carefully placed predominantly within the gastrocnemius
medialis and lateralis muscles (Fig
1). Water was shimmed
to 35–45 Hz (magnitude spectra).
Dynamic semi-LASER-localized3,4 1H MR spectra (TE=30ms) (Cr) and DRESS-slab-localized5 31P MR spectra (PCr, Pi) were acquired
interleaved in one exercise/recovery session (3min rest, 5min exercise at 30% of maximal voluntary
contraction force, 10min recovery) with
a TR of 6 s, similarly as published6. Volunteers were pushing the pedal twice per TR, audio-cued
by gradient, to ensure that data were acquired always in relaxed muscle.
1H/31P
spectroscopy data were processed from raw data using in-house-developed Python
scripts (http://www.python.org), phasing signals to the highest peak magnitude
of PCr, water, or lipids in the frequency domain for channel combination. Peak amplitudes were quantified with AMARES, using
jMRUI v6.0 alpha. For fitting the 3.9 ppm Cr-CH2 resonance, we used HLSVD
peak removal over the lipid resonances at 1.5 ppm (HLSVD no-max-2048-points
filter, 5 components).
For
absolute quantification of the visible fraction of Cr-CH₂ resonance, the fully-relaxed
water signal was measured separately. Concentrations were calculated in mmol/L (tissue volume) units, by: Cm=CH2O x (Sm/SH2O) x (nH2O/nm) x (CFH2O/ CFm) x WH2O x
ρmuscle
with S the signal intensity of water or Cr, nH2O
and nm the number of equivalent protons (= 2), CF the
correction factors for T1 and T2 relaxation, CH2O =
55.56 mol/L the
water concentration, CFm the approximate water content in skeletal muscle
tissue, (0.77 L/kgww), and ρmuscle the specific weight of
skeletal muscle tissue (1.06 kg/L).
31P MRS yielded PCr and Pi concentrations, relative
PCr depletion and depletion-rate during exercise (τPCr on-kinetics)
and resynthesis-rate during recovery (τPCr
recovery), the
maximal oxidative phosphorylation rate, i.e., mitochondrial capacity (Qmax),
and the time course of intracellular pH7,8
.
Data are presented
as mean ± standard deviation. Results
Submaximal
exercise leads to disappearance of the Cr-CH2 resonance at 3.92ppm
in 1H MR spectra (Fig 2, Fig 3) whereas the Cr-CH3
resonance at 3.02ppm remained stable during exercise. Detailed numerical results
regarding concentrations
and time constants are shown in Table 1, individual data in Fig 4.
Most strikingly, the mean time constants of PCr and Cr-CH2 recovery dynamics were not
significantly different, τPCr = 37±9 s and τCr = 34±6 s, p=0.2, showing marginally positive
correlation (r=0.69, p=0.058). τPCr on-kinetics and τCr-CH2 on-kinetics were positively correlated (r=0.73, p=0.039). Discussion and Conclusion
We assessed the skeletal muscle Cr-CH2 resonance together
with PCr during a single dynamic session using interleaved 1H/31P
MRS at 7T. Despite limited visibility of the Cr-CH2 resonance, we
were able to quantify both time courses in a single
experiment in eight subjects. Therefore, we were able to directly compare Cr-CH2
and PCr time courses during exercise and recovery. We observed remarkably similar
kinetics of Cr-CH2 and PCr time courses in absolute numbers, and
positive correlations between τPCr and τCr-CH2. The discrepancy between end-exercise depletion
measured with 1H and 31P MRS may be explained by
differing volumes of interest9. Studies
including higher numbers of subjects will be necessary to explain quantitative
differences, especially found in exercise-on kinetics, and verification
assessing skeletal muscle oxidative metabolism by 1H MRS.Acknowledgements
Colleagues at the
High-Field MR Center, this study was supported by FWF (#KLI 904 to MKrss)References
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