A Ankeeta1, S Senthil Kumaran1, and Rohit Saxena2
1Department of NMR & MRI Facility, All India Institute of Medical Sciences, Delhi, India, 2Dr RP Centre of Ophthalmology, All India Institute of Medical Sciences, Delhi, India
Synopsis
This study investigated structural alteration, cerebellar BOLD activation
and functional connectivity during Braille reading by congenital blind (CB)
children. Functional magnetic resonance imaging (fMRI) during Braille reading and
T1 data were acquired on a 3T MR scanner in CB and sighted control (SC)
children. CB children showed bilateral activation in posterior lateral
cerebellum Crus I, II. BOLD cerebellar, visual signals and functional
connectivity measures exhibited positive correlation with duration of Braille
reading. Involvement of cerebellum, visual cortex during Braille reading in congenital
blind children suggest its role in haptic language processing.
Background
Non-visual perception in visually impaired subjects
require interaction between attention and sensory systems to infer the stimuli
and its spatial location. Mode of stimuli input (Braille/Auditory) induces difference in language
processing, spatial and feature-based attention networks after visual deficit.1 Involvement
of visual cortex and an increased involvement of cerebellum for non-visual
stimulation in congenitally blind subjects (than sighted control) has been
associated with perception, language and attention.2 This study explores
cortico-cerebellar functional connectivity for non-visual processing of
language components over other sensory systems in CB children.Methodology
High-resolution T1W images and task based functional MR
images of 20 congenital and 15 sighted control subjects
(Table 1) were acquired using 3T MR scanner (Achieva 3T TX). Braille reading
task comprised of 16 nouns (followed by four synonym options for each word),
presented in block design, with Blood oxygenation level dependent (BOLD) signal
measurement in 192 dynamics (TR/TE: 2000/30ms, 29 contiguous slices, 4.5mm
thickness). The structural changes of cerebellum were evaluated using SUIT toolbox3
and Individual Component Analysis using CONN toolbox.4,5 Influence
of language and cerebellum areas on task-based fMRI was investigated using
three connectivity measures (global connectivity, local connectivity and
cluster coefficient task). Correlation of functional connectivity measure were
done with duration of Braille reading.Results
Seed to target analysis for ROI analysis exhibit
increased FC between the posterior cerebellar (pCP), visual and language
networks (LN). Differences in FC associated with duration of exposure of
language semantic were observed in visual (VN), cerebellar (CP; CA), and
language components (Table 2, Figure 1). Increased BOLD activation and gray matter volume were observed in cerebellar lobule
in Vermis 3, lateral portion of Crus I and II and posterior cerebellar areas of
CB compared to SC. A positive correlation of BOLD and FC measures with duration
of Braille reading
was observed in congenital blind.Discussion
Increased functional connectivity was
observed between language regions (inferior frontal gyrus, superior temporal
gyrus, middle temporal gyrus, inferior temporal cortices, precuneus) and bilateral
cerebellar areas (crus I, crus II, cerebellum 45). Functional involvement of
visual cortex, IFG, STG, MTG and cerebellum during
braille reading task may be attributed to language processing.1,2 Functional
connectivity of cerebellum with visual cortex, prefrontal, MTG and hippocampus in
congenital blind group exhibit intact language perception and processing
networks and increased correlation with duration of Braille reading may be attributed
to non-visual processing involving comprehension of complex semantics and
encoding of concrete words.1,6,7
Considering the tactile based Braille reading and enhanced connectivity
involving the multi-sensory regions, results reflect a compensatory mechanism in
congenital blind.Conclusion
Increased strength of connectivity and BOLD activation in
cerebellum suggest intact language processing through fluency and decoding of
haptic language.Acknowledgements
The authors acknowledge Department of Science and technology, Government of India for funding the study. References
1. Amedi A, Malach R, Hendler T, Peled S, Zohary E.
Visuo-haptic object-related activation in the ventral visual pathway. Nat
Neurosci. 2001;4(3):324–30.
2. Buchel C. Functional neuroimaging studies of
Braille reading: cross-modal reorganization and its implications. Brain.
1998;121(7):1193–4.
3. Diedrichsen, J. et al. Imaging the Deep
Cerebellar Nuclei: A Probabilistic Atlas and Normalization Procedure.
NeuroImage. 2011;54(3): 1786–94.
4. Whitfield-Gabrieli S, Nieto-Castanon A. Conn : A
Functional Connectivity Toolbox for Correlated and Anticorrelated Brain
Networks. Brain Connect. 2012;2(3):125–41.
5. Friston KJ, Holmes AP, Price CJ, Büchel C, Worsley KJ.
Multisubject fMRI studies and conjunction analyses. Neuroimage. 1999;10(4):385–96.
6. Stoodley,
C. J., & Schmahmann, J. D. The cerebellum and language: evidence from patients
with cerebellar degeneration. Brain and Language. 2009;10(3), 149–53.
7. Wallentin M, Ostergaard
S, Lund TE, Ostergaard L, Roepstorff A: Concrete spatial language: see what I
mean? Brain Lang 2005, 92: 221–233.