Anirban Sengupta1, Arabinda Mishra1, Feng Wang1, Li Min Chen1, and John C. Gore1
1Vanderbilt University Medical Center, Nashville, TN, United States
Synopsis
The
objective of this study was to identify fine-scale resting state functional
networks within the spinal cord gray matter of squirrel monkeys, and measure
the changes in functional connectivity within the cord after a targeted injury.
Independent Component Analysis of resting state fMRI data detected
robust BOLD signals localized at the bilateral intermediate and gray-commissure
regions of the spinal cord as well at the ‘4 horns’. A unilateral section of
dorsal column tract at C5 segment of spinal cord damaged the inter-segmental
connectivity more than intra-segmental connectivity, as observed through
individual connectivity measures and community structures generated by
graph-theory principles.
INTRODUCTION
Robust
measures of functional connectivity based on correlations between blood-oxygenation-level-dependent
(BOLD) signals in a resting state have been derived from gray matter (GM)
within the spinal cord (SC) in several studies recently1-3. However,
connectivity in SC has been interpreted in terms of relatively simple models,
mainly confined to the four dorsal and ventral horns which are known to be
responsible for transmission of sensory and motor signals to and
from the brain2-4. We sought to extend our knowledge of the functional
networks within SC GM beyond the ‘4 horn model’ by adopting a data-driven
method of analysis that does not require a priori identification of seed
regions of interest (ROI). Independent Component Analysis (ICA) was used to
identify clusters of voxels that show temporal synchrony in BOLD fluctuations
independent of other clusters, within the gray matter of the SC of squirrel
monkeys and these were used as seed locations for subsequent evaluations of connectivity
within/intra and between/inter spinal segments5. Changes in network
organization and connectivity at intra and inter-segmental level were also
measured after a targeted injury to the spine, at various stages of recovery.METHODS
Images of five
axial slices, covering C3-C7 cervical
segments of anaesthetized squirrel monkeys were acquired using a custom neck
coil by an Agilent 9.4T scanner. Resting state fMRI data (300 dynamics,14 runs
from 5 monkeys) were acquired using a fast gradient echo sequence (flip angle =
~18°, TR = 46.88ms, TE: 6.5ms, 3s per volume). Data were acquired before and
after a unilateral surgical transection of dorsal column at C5 level (Fig1).
Post SC injury (SCI), data were collected at three time-points: after 2weeks,
7-8weeks and 16-22weeks. Motion, physiological signal correction (RETROICOR) and
band pass filtering (0.01-0.1 Hz) were performed on the fMRI data, followed by
co-registration to a customized template using FSL to facilitate group level analyses6,7.
Group spatial ICA was performed by temporal concatenation of all the data using
GIFT software, and thirty-five spatially independent components were extracted
within the butterfly-shaped gray-matter region of SC5. Next,
thirty-four ROIs were identified across the five spine segments by visual
inspection of each component’s spatial profile. Connectivity was measured between
ROI pairs as the correlation of time courses of IC pairs. Connectivity values
between FDR corrected significant ROIs (p<0.05) were computed before and at different
time points after injury, followed by statistical tests for comparison. Next,
graph-theory analysis was performed to generate community structures based on
connectivity measures, which depicted functional network organization within
and across the spinal segments8. Finally, the connectivity matrix
was rearranged such that the connectivities within each community
are next to each other to illustrate the community behavior.RESULTS
ICA
decomposed resting state fMRI data into seven spatially distinct ROIs in each
spinal segment: bilateral dorsal horns, bilateral ventral horns,
bilateral intermediate region which extended
from the dorsal to the middle part of SC, and gray-commissure
region, which is the thin
strip of gray matter located at the central junction between the two hemi-cords, as
shown for a representative slice in Fig2A. Significant intra-slice/intra-segment
and neighboring inter-slice/inter-segment connectivities were noted and used
for further analysis as depicted by the connectivity matrix (Fig2B). Fig3C shows
that post-SCI there was a significant (p<0.05) drop in connectivity at
Stage1, for all the neighboring inter-slice connectivities, whereas this was
true for only Slices 2 & 3 for intra-slice connectivities (Fig3B). There
was a gradual increase in both intra-slice and neighboring inter-slice
connectivity over time after injury with all of them recovering back to normal
values by the last time point (except Slice3-Slice4 neighboring slice
connectivity). The communities formed from graph-theory depicted higher
intra-community and lower inter-community connectivity as evident visually from
the connectivity values of the rearranged connectivity matrix (Fig4A). Before
injury, the community structure was spread within and across multiple slices/segments
(Fig4B). Post SCI at Stages 1&2 the communities become confined to ROIs
within each slice before starting to reappear at the inter-segment level by the
last time point.DISCUSSION
Data-driven ICA analysis observed the presence
of localized BOLD fMRI coherent fluctuations at the bilateral intermediate
regions of cervical SC of monkeys which agrees with our recent findings of similar
activity observed using electrophysiology9. The presence of strong localized connectivity at the gray
commissural region possibly mediates communications between two sides of SC9,10.
Targeted dorsal column lesion had a differential effect on intra-slice and neighboring inter-slice
connectivity with the reduction in connectivity being more
evident in the latter. This finding was also supported by graph-theory results
which found that community structures were reduced to intra-segments post
injury. Overall, graph-theory results suggest that SCI primarily damages inter-segment
community formation which slowly reverts to baseline structure with time.CONCLUSION
This study highlights the presence of robust,
correlated BOLD signals at the bilateral intermediate regions and gray-commissure
regions of cervical spinal cord, in addition to the dorsal and ventral horns.
This indicates the presence of a complex fine-scale intrinsic functional
network in SC. A targeted injury damages inter-segment community formation and reduces
connectivity primarily between the segments than within them. The connectivity
measures returned to baseline by the last time point after injury which
signifies recovery.Acknowledgements
This study is supported by NIH grant NS092961 and DOD grant SC160154.References
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