Emma Biondetti1,2,3, Mathieu D. Santin1,2, Romain Valabrègue1,2, Graziella Mangone1,4, Rahul Gaurav1,2,3, Nadya Pyatigorskaya1,3,5, Matthew Hutchison6, Lydia Yahia-Cherif1,2, Nicolas Villain1,7, Marie-Odile Habert8, Isabelle Arnulf1,3,9, Smaranda Leu-Semenescu9, Pauline Dodet9, Jean-Christophe Corvol1,4,7, Marie Vidailhet1,3,7, and Stéphane Lehéricy1,2,3,5
1Institut du Cerveau – ICM, INSERM U 1127, CNRS UMR 7225, Sorbonne Université, Paris, France, 2ICM, Centre de NeuroImagerie de Recherche – CENIR, Paris, France, 3ICM, Team “Movement Investigations and Therapeutics” (MOV’IT), Paris, France, 4National Institute of Health and Medical Research - INSERM, Clinical Investigation Centre, Pitié-Salpêtrière Hospital, Paris, France, 5Department of Neuroradiology, Pitié-Salpêtrière Hospital, AP-HP, Paris, France, 6Biogen Inc., Cambridge, MA, United States, 7Department of Neurology, Pitié-Salpêtrière Hospital, AP-HP, Paris, France, 8Department of Nuclear Medicine, Pitié-Salpêtrière Hospital, AP-HP, Paris, France, 9National Reference Center for Rare Hypersomnias, Pitié-Salpêtrière Hospital, AP-HP, Paris, France
Synopsis
Parkinson's disease (PD) and idiopathic rapid eye movement
sleep behaviour disorder (iRBD, a prodromal condition of Parkinsonism) are
characterised by progressive striatal dopaminergic denervation, loss of
neuromelanin-containing neurons and increased iron deposition in the substantia
nigra. Here, we evaluated neurodegeneration-induced changes in the nigrostriatal
system of PD and iRBD patients using longitudinal SPECT and MR imaging compared
to healthy control subjects. We showed that dopamine, neuromelanin and iron
changes followed similar spatiotemporal gradients of neurodegeneration, and we assessed
the temporal onset and ordering of such changes.
Introduction
Parkinson's disease (PD) is diagnosed when 30-50%
dopaminergic neurons in the substantia nigra pars compacta (SNc) are lost.1,2
Motor symptoms in PD are often preceded by early non-motor symptoms, e.g.
idiopathic rapid eye movement sleep behaviour disorder (iRBD).3
Neuromelanin (NM) and iron are interrelated via the dopaminergic
metabolic pathway and may participate in the neurodegenerative process of PD.4,5
Indeed, the NM-containing neurons of PD patients have abnormally elevated iron
concentrations, up to 50% higher than in healthy controls (HCs).6,7
Previous studies using a combination of NM-sensitive MRI
(NM-MRI), iron-sensitive MRI, and dopamine-sensitive nuclear imaging have correlated
striatal dopamine decrease and SNc NM decrease,8-11 striatal dopamine
decrease and SNc iron increase,11 SNc NM decrease and SNc iron
increase.12-15 These three measurements have never been investigated
together longitudinally in both clinical PD and prodromal PD (iRBD). Other
studies have temporally placed the onset of striatal dopamine changes (10-25
years before PD diagnosis)16 or nigral NM (5 years before PD diagnosis).17
However, the time onset of nigral iron changes is unknown.
Here, we aimed to investigate the spatiotemporal relationships
between iron, NM and dopamine transporter (DaT) in the nigrostriatal system of
patients with PD and iRBD.Methods
A longitudinal cohort was prospectively investigated,
including (#visit 1[#visit 2]): 55[28] HCs, 43[21] iRBDs and 135[83] PDs (Table
1). Motor disability, global cognition and mood/behaviour were respectively
evaluated using the MDS-UPDRS part III;18 the Mattis DRS19,20
and the MoCA scales;21 and the ASBPD scales22 (Table 1).
MRI was performed on a Siemens Prisma 3 T system (64-channel
receive head coil) including a 3D T1-weighted MP2RAGE acquisition
for anatomical reference, a 2D T1-weighted TSE protocol for NM-MRI,
and a 3D T2*-weighted FLASH protocol for iron-sensitive MRI (Table
2). 35[18] HCs, 32[16] iRBDs, 48[40] PDs also underwent DaT-SPECT on a hybrid gamma
camera Discovery 670 Pro system using the 123I-Ioflupane tracer.
For each subject, a quantitative susceptibility map (QSM)
was calculated based on the iron-sensitive data (QSM pipeline: nonlinear temporal
fitting,23 background field removal using projection onto dipole
fields,24 inverse problem solution using piece-wise constant
regularization,25 referencing to the posterior limb of the internal
capsule26). Both the NM-sensitive and QSM images were coregistered
to a digital brain template as previously described.17 In template
space, an SNc region of interest (ROI) previously calculated based on NM-MRI of
HCs17 was considered, and here voxel-wise values of NM signal to
noise ratio (SNR) and QSM were calculated. The SNc ROI was manually subdivided
into three subregions:27 sensorimotor, associative and limbic. In
each SNc subregion, the average NM SNR and QSM signal were calculated. For each
subject, the DaT-SPECT was aligned to the MP2RAGE image, then referenced to an
ROI in the occipital cortex with non-specific DaT uptake.28 The
average DaT (specific binding ratio) was calculated in seven striatal ROIs segmented
by aligning the YeB atlas29,30 to the MP2RAGE image: the nucleus
accumbens (NAc), the sensorimotor, associative and limbic caudate nucleus (Cd)
and the sensorimotor, associative and limbic putamen (Pu). In patients, all NM,
QSM and DaT values were lateralized to obtain consistent alignment with the
most affected brain hemisphere.
In the most affected hemisphere, the temporal
evolution of striatal DaT, SNc NM and SNc iron was investigated by exponentially
fitting their percent ratio relative to HCs over disease duration. Inter-group
differences in striatal DaT, nigral NM and nigral iron at V1, V2 and
longitudinally were assessed using univariate linear models (covariates: sex
and age, Tukey’s correction for multiple comparisons). In the SNc, concomitant
changes between striatal DaT, SNc NM and SNc iron were evaluated by calculating
voxel-wise correlations between each pair of measurements (adjusted for
multiple comparisons).Results and Discussion
In the striatum and SNc of patients, based on the
inter-group differences in DaT, NM and iron, neurodegeneration appeared to
start in the most affected hemisphere (progressing from sensorimotor, to
associative and limbic areas) and to progress with a similar spatial pattern
and a lower extent in the contralateral hemisphere (Figure 1).
In the most
affected hemisphere of PDs, the analysis of DaT, NM and iron over disease
duration indicated that DaT loss started between 20.7 (sensorimotor Pu) and 1.9
years (limbic Cd) before PD diagnosis (Figure 2). In the SNc, iron accumulation
started 9.6 (sensorimotor), 6.2 (associative) or 4.1 years (limbic) before
diagnosis, whereas NM loss started 5.9 (sensorimotor) and 3.6 years (associative)
before diagnosis (Figure 2). In the limbic SNc, NM signal changes were very
small.
In the SNc, the spatial
DaT-NM correlation pattern was lateralized in the most affected hemisphere,
whereas the DaT-iron correlation pattern was always bilateral (except for the NAc)
(Figure 3). The NM-iron correlation pattern was located in the anterior SNc,
corresponding to associative/limbic territories (Figure 3).
Taken
together, these results suggest that in PD, first DaT is lost in the striatum,
then iron accumulation precedes NM loss in the SNc. All three measurements were
correlated.Conclusions
In the nigrostriatal system in prodromal and clinical PD, we
showed the spatial gradient of changes in DaT, NM and iron, and estimated the
temporal onset and the relative temporal ordering of such changes. DaT, NM and
iron were correlated suggesting a relationship between iron changes and
degeneration of the dopaminergic pathway.Acknowledgements
This study was funded by grants from the Investissements
d'Avenir, IAIHU-06 (Paris Institute of Neurosciences – IHU), ANR-11-INBS-0006,
Fondation d’Entreprise EDF, Biogen Inc., Fondation Thérèse and René Planiol,
Fondation Saint Michel, Unrestricted support for Research on Parkinson’s disease
from Energipole (M. Mallart) and Société Française de Médecine Esthétique (M.
Legrand). Emma Biondetti received grant funding from France Parkinson and Biogen Inc. The
authors would like to thank all the subjects who participated in this study.References
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