Deanne K Thompson1,2,3,4, Joseph YM Yang2,3,5,6, Jian Chen2, Claire E Kelly1,2, Chris L Adamson2, Bonnie Alexander1,2, Lillian G Matthews7, Katherine J Lee1,3,8, Rod W Hunt1,3,9, Jeanie LY Cheong1,10,11, Megan Spencer-Smith1,12, Marc L Seal2,3, Terrie E Inder1,7, Lex W Doyle1,3,10,11, and Peter J Anderson1,12
1Victorian Infant Brain Studies, Murdoch Children's Research Institute, Parkville, Australia, 2Developmental Imaging, Murdoch Children's Research Institute, Parkville, Australia, 3Department of Paediatrics, University of Melbourne, Parkville, Australia, 4Florey Institute of Neuroscience and Mental Health, Parkville, Australia, 5Neuroscience Research, Murdoch Children’s Research Institute, Parkville, Australia, 6Department of Neurosurgery, the Royal Children’s Hospital, Parkville, Australia, 7Department of Pediatric Newborn Medicine, Brigham and Women’s Hospital, Harvard Medical School, Boston, MA, United States, 8Clinical Epidemiology & Biostatistics Unit, Murdoch Children’s Research Institute, Parkville, Australia, 9Neonatal Medicine, Royal Children’s Hospital, Parkville, Australia, 10Royal Women's Hospital, Parkville, Australia, 11Department of Obstetrics and Gynaecology, University of Melbourne, Parkville, Australia, 12Turner Institute for Brain and Mental Health, School of Psychological Sciences, Monash University, Clayton, Australia
Synopsis
Children born very preterm have altered
myelination compared with full-term controls, but whether the T1-T2 ratio is a sensitive measure for
understanding neurodevelopmental functioning remains unknown. This study found
that the T1-T2 ratio trajectory
between 7 and 13 years of age in the uncinate fasciculus was related to IQ
scores, and 13-year T1-T2 ratios in
almost all white matter regions were associated with motor functioning in both
birth groups. Myelin
development assessed using the T1-T2 ratio appears to be sensitive to predicting some neurodevelopmental outcomes.
Introduction
Myelination begins late in fetal development, and
continues into early adulthood.(1) Myelin mapping, using the T1-weighted and T2-weighted
structural MRI images as a ratio, may provide insight into the maturity of
myelin.(2,3) Children born very preterm (VP) are known to have
developmental vulnerability of myelin.(4)
We have recently shown through a
longitudinal study that VP children at 7 and 13 years of age have reduced T1-T2 ratios, predominantly in the white matter of the
limbic system, compared with full-term (FT) children.(5) We hypothesise that this may explain some of the
cognitive, learning, memory and behavioral problems VP children face.(6) We have also shown that VP children born small for
gestational age have reduced T1-T2 ratios predominantly in
the sensorimotor tracts,(5) and hypothesise that this may explain some of their
reported motor problems.(7) However, it is unknown whether the T1-T2 ratio is related to these neurodevelopmental
outcomes.
The aims of this study were to
1) examine
associations between (a) the T1-T2 ratio at
term-equivalent age, 7 or 13 years of age, or (b) the trajectories of T1-T2
ratio development between term and 13 years, and cognitive,
learning and memory, behavioral and motor functioning at 13 years, and 2)
assess whether these associations differed between the VP and FT groups.Methods
VP children born <30 weeks’ gestation or
very low birthweight (<1250 g) and FT controls (born ≥37 weeks’) underwent longitudinal T1
and T2-weighted brain MRI. 224 VP and 45 FT infants were scanned at term-equivalent age, 159 VP and 36 FT children at 7 years, and 140 VP and
48 FT children at 13 years corrected age, with 198 VP and 56 FT children having usable T1 and T2
scans at term-equivalent, 7 and/or 13 years of age.
T1-
and T2-weighted images were first corrected for β1 bias field inhomogeneity(8) and brain extracted.(9) T1-T2 ratio whole brain myelin map images were then
calculated. The myelin maps were parcellated into white matter regions of
interest using the JHU neonatal(10) and adult(11) atlases. 47 brain regions were assessed across the 3
time-points.
Neurodevelopmental outcomes were collected
at 13 years of age for intelligence quotient (IQ) using the Kaufman Brief
Intelligence Test, Second Edition (KBIT-2), motor outcomes using the Movement
Assessment Battery for Children, Second Edition (MABC-2), memory and learning
using the California Verbal Learning Test, Children’s Version (CVLT-C), and
behavior using the Strengths & Difficulties Questionnaire (SDQ).
Linear regression models were used to examine associations between T1-T2 ratios at
each time-point, the rate of change in T1-T2
ratios from term-equivalent to 7 years and 7 to 13 years, and neurodevelopmental
outcomes. A group-by-T1-T2 ratio interaction term was included, and all estimates were adjusted for sex,
age at assessment, and social risk at 13 years. P-values were False
Discovery Rate corrected.Results
Increases in T1-T2 ratio between 7 and 13 years in the left uncinate fasciculus were
associated with higher IQ (p=0.025), across both birth groups.
Higher T1-T2 ratios in all brain
regions at 13 years of age were associated with better motor functioning (all p<0.026),
except the fornix column and body, and superior cerebellar peduncles.
There were no associations between the T1-T2 ratio at or across any time-points with memory and
learning or behavioral scores (all p>0.07).Discussion and conclusion
In line with our hypotheses, we found that
increasing T1-T2 ratio
trajectories between 7 and 13 years in one of the limbic structures, the
uncinate fasciculus, were associated with higher IQ at 13 years of age in both VP and FT children. This suggests
that our previous findings of reduced myelin content in the limbic structures
of VP children compared with FT children may have consequences for cognitive
functioning.
Higher T1-T2 ratios at 13 years were
also associated with better concurrent motor functioning, with no regional or
birth group specificity. However,
early measures of T1-T2 ratio at term-equivalent
age did not appear to be sensitive to later functional outcomes.
Myelin development assessed using the T1-T2 ratio may help to explain poorer IQ and motor outcomes experienced
by VP children at 13 years of age. Acknowledgements
We would like to thank members of the
Victorian Infant Brain Studies and Developmental Imaging groups the at the
Murdoch Children Research Institute for ideas and support, Michael Kean and
Radiographers at the Royal Children’s Hospital for the magnetic resonance
images, and the families and children who participated in this study.
This study was supported by the Australian
National Health and Medical Research Council (NHMRC) (Centre for Clinical
Research Excellence 546519; Centre for Research Excellence 1060733; Project
Grants 237117, 491209 & 1066555; Senior Research Fellowships 628371 &
1081288 to PJA; Career Development Fellowship 1085754 to DKT; Early Career
Fellowship 1012236 to DKT), US National Institutes of Health HD058056, United
Cerebral Palsy Foundation (USA), Leila Y. Mathers Charitable Foundation (USA),
the Brown Foundation (USA), Murdoch Children’s Research Institute, the Royal
Children’s Hospital, The Royal Children’s Hospital Foundation, Department of
Paediatrics, The University of Melbourne and the Victorian Government's
Operational Infrastructure Support Program.
References
1. Yakovlev PI, Lecours A. The
myelogenetic cycles of regional maturation of the brain. In: Minkowski A,
editor. Regional Development of the Brain in Early Life. Oxford: Blackwell
Science; 1967.
2. Glasser MF, Van
Essen DC. Mapping human cortical areas in vivo based on myelin content as
revealed by T1- and T2-weighted MRI. J Neurosci 2011;31(32):11597-11616.
3. Ganzetti M,
Wenderoth N, Mantini D. Whole brain myelin mapping using T1- and T2-weighted MR
imaging data. Front Hum Neurosci 2014;8:671.
4. Volpe JJ, Kinney
HC, Jensen FE, Rosenberg PA. The developing oligodendrocyte: key cellular
target in brain injury in the premature infant. Int J Dev Neurosci
2011;29(4):423-440.
5. Thompson DK, Yang
J, Chen J, et al. Longitudinal Myelin Development in Children Born Very Preterm
Compared with Typically Developing Peers. International Society for Magnetic
Resonance in Imaging Annual Meeting. Paris, France; 2018.
6. Treyvaud K, Doyle
LW, Lee KJ, et al. Social-emotional difficulties in very preterm and term 2
year olds predict specific social-emotional problems at the age of 5 years. J
Pediatr Psychol 2012;37(7):779-785.
7. Murray E,
Fernandes M, Fazel M, Kennedy SH, Villar J, Stein A. Differential effect of
intrauterine growth restriction on childhood neurodevelopment: a systematic
review. BJOG : an international journal of obstetrics and gynaecology
2015;122(8):1062-1072.
8. Tustison NJ,
Avants BB, Cook PA, et al. N4ITK: improved N3 bias correction. IEEE Trans Med
Imaging 2010;29(6):1310-1320.
9. Smith SM. Fast
robust automated brain extraction. Hum Brain Mapp 2002;17(3):143-155.
10. Oishi K, Mori S,
Donohue PK, et al. Multi-contrast human neonatal brain atlas: Application to
normal neonate development analysis. NeuroImage 2011;56(1):8-20.
11. Oishi K, Zilles K,
Amunts K, et al. Human brain white matter atlas: identification and assignment
of common anatomical structures in superficial white matter. Neuroimage
2008;43(3):447-457.