Choong Heon Lee1, Jing Li2, Yulin Ge1, Timothy M Shepherd1, Youssef Zaim Wadghiri1, Jiangyang Zhang1, and David W Nauen3
1Radiology, New York University School of Medicine, New York, NY, United States, 2Peking Union Medical College Hospital, Beijing, China, 3Pathology, Johns Hopkins University School of Medicine, Baltimore, MD, United States
Synopsis
High-resolution diffusion MRI data
of post-mortem adult human hippocampus specimens were acquired and compared to
histology to identify major axonal pathways in the hippocampus. The complex
microstructural organization in the hippocampus made it difficult to resolve
axonal pathways based on conventional diffusion tensor data. In comparison,
neurite density map using the NODDI toolbox revealed the locations of the
perforant path, mossy fibers, and Schaffer collaterals confirmed by histology.
We were able to reconstruct the fimbria/alveus and perforant pathways using
tractography, and the results resembled in
vivo results from the HCP dataset. Other pathways in the hippocampus
remained difficult to delineate.
Introduction
The hippocampus plays an important role in
memory and learning1. It
consists of several subregions, which have unique cyto-architectural features. Axonal
pathways including the perforant path, mossy fibers, and Schaffer collaterals convey
information among these subfields and other brain regions, mainly the
entorhinal cortex. It has long been postulated that the morphology and microstructural
organization of hippocampal areas and the structural integrity of the
hippocampal pathways may serve as markers of present or future memory ability.
While software tools have been developed for segmentation of hippocampal
subfields2,3, and diffusion MRI (dMRI) tractography has been used to
trace some hippocampal pathways4 and correlate dMRI signals with
memory function5, visualization of microstructural organization,
especially that of axonal pathways, within the hippocampal subfields remains limited. Several studies have used post-mortem structural and diffusion
MRI to examine the human hippocampus at high spatial resolution 6-10.
In this study, we combined high-resolution diffusion MRI and histology to
examine neural pathways in the hippocampus. Methods
The
study was approved by the Institutional Review Board. De-identified post-mortem
specimens of formalin-fixed adult human hippocampus (age 16-44 years, n=3) were
obtained after pathological autopsy examination following family consent. Experiments
were performed on a horizontal 7-T MRI system (max gradient strength=600mT/m)
using a 30 mm diameter volume transmit/receive coil at room temperature with
the following parameters: 1) 2D multi-shell dMRI: 4-shot EPI, TE/TR = 38.2/6000
ms, δ/Δ = 6/24
ms, BW=200KHz, 60 diffusion directions with 6 b shells from 2,000 to 12,000
s/mm2; 2) 3D single-shell dMRI: 3D diffusion-weighted GRASE, TE/TR =
40/500 ms, d/D =
6/20 ms, 0.25 mm isotropic resolution, BW=200KHz, 60 diffusion directions, b = 5000
s/mm2; 3) 3D T2-weighted MRI: RARE, TE/TR = 50/2000ms, ETL=8, 0.25 mm isotropic resolution. dMRI data were analyzed using MRtrix and NODDI11
toolboxes. Tissue sections were stained with hematoxylin and eosin (H&E,
cellular structure), neurofilament (EMD Millipore, clone SMI-31, axons), and Hirano
silver (cell bodies and processes).Results
Normalized
diffusion-weighted signal attenuation curves (Fig.1) show that gray matter had
faster signal decays than white matter, but the contrast mostly stabilized for diffusion
weighting (b) greater than 5,000 s/mm2. Results from fitting the 2D
multi-shell dMRI data to diffusion tensor imaging (DTI) and neurite orientation
dispersion and density imaging (NODDI) models were compared with histology at
similar locations (Fig. 2). Based on the DTI, histology, and previous reports,
we segmented hippocampal subfields in the 3D dMRI volume (Fig. 2). The dentate
gyrus had higher mean diffusivity (MD) values than surrounding regions. The CA1
region (Fig. 3, white arrows) showed moderate fractional anisotropy (FA) values
and consistent radial orientation, likely due to coherently arranged apical
dendrites in this region, as shown by silver staining.
The
neurite density index (NDI) and orientation dispersion index (ODI) maps
revealed additional information on hippocampal pathways. The NDI map showed
several neurite areas in the CA1-CA3 subregions (Fig. 3, red, blue, and green
arrows), in good agreement with SM31 stained sections showing axons in the
perforant path (blue), Schaffer collaterals (red), and mossy fibers (green).
The same regions had low diffusion anisotropy and are difficult to visualize
using DTI, probably due to crossing of these axons with apical dendrites of the
pyramidal neurons. This is supported by high orientation dispersion in these
regions shown by the ODI map.
Using fiber orientation
distribution (FOD) maps (Fig. 4A) estimated from the 3D dMRI volume and
subfield segmentation (Fig. 2), reconstruction of the fimbria/alveus (Fig. 4B),
perforant pathway (Fig. 4C), and mossy fiber pathway
(Fig. 4D) were compared to in vivo
results from the Human Connectome Project data (not shown here). Our results
suggest that fimbria/alveus and perforant pathways can be reconstructed from
dMRI data but the mossy fiber and other intrahippocampal pathways remain challenging
to reconstruct due to the large population of
orthogonally-oriented apical dendrites. Discussions
Our
results demonstrate that NODDI can reveal additional microstructural
information related to neural pathways in the human hippocampus. Previous
reports have described similar findings in hippocampal subfields10.
The histological data reported in this study further confirms the NODDI
results. It is necessary to note that NODDI results depend on initial setting
of intra and extracellular diffusivities.
Our
results also suggest that several intra-hippocampal pathways are difficult to
reconstruct using conventional dMRI tractography tools due to the presence of
apical dendrites. The limitations in identifying these structures can also be
attributed to tissue degradation after death. However, little degradation was
noted in the current case based on the corresponding histological sections.
This limitation raises the need to further refine the tractography tools
utilizing additional information provided by NODDI and other tissue
microstructural models. Conclusion
Despite
improvement on the spatial resolution at
0.25 mm, some of the hippocampal pathways remained difficult to reconstruct.
However, NODDI proved very useful in providing additional information to
delineate neural pathways in the hippocampus.Acknowledgements
No acknowledgement found.References
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