Fabian Niess1,2, Albrecht Ingo Schmid1,2, Siegfried Trattnig2,3, Ewald Moser1,2, and Martin Meyerspeer1,2
1Center for Medical Physics and Biomedical Engineering, Medical University of Vienna, Wien, Austria, 2High-Field MR Centre, Medical University of Vienna, Wien, Austria, 3Department of Biomedical Imaging and Image-guided Therapy, Medical University of Vienna, Wien, Austria
Synopsis
The purpose of this study is to show the effect of low-intensity
exercise on localised 31P MR spectra and 1H
images of human muscle. It was determined which PCr depletion is
sufficient for quantifying the PCr recovery time constant (τPCr),
while incurring minimal pH drop in a predominantly glycolytic muscle.
PCr depletion during exercise was measured, PCr recovery was fitted
and pH was quantified with dynamic localised 31P MRS, while
perfusion and BOLD 1H images were simultaneously acquired in time-resolved measurements. Prolongation of τPCr
with acidification was confirmed, while very short τPCr
was found with neutral and slightly acidic pH.
Introduction
Exercise/recovery studies of human muscle using 31P MRS have mostly
been performed by prescribing a fixed exercise intensity as a
fraction of the subjects’ maximum voluntary contraction force
(MVC), typically 30 % or more. The purpose is to deplete
phosphocreatine (PCr) sufficiently much to measure the time constant
of its recovery post-exercise, τPCr. This rate is
closely tied to the oxidative ATP production rate, and is hence used
to quantify muscle ‘mitochondrial capacity’ 1 –
which is straight-forward only if pH at end of exercise (pHee)
does not fall by more than 0.1 – 0.2 units due to glycolytic
contributions to ATP synthesis. While quite significant PCr depletion
(~ 50 %) has been reached without observations of acidification with
in partly- or unlocalised MRS 2, more recent experience
with muscle-specific 31P MRS localisation reflected pH changes at
already relatively low exercise intensities 3,4. A way to
avoid acidification is to monitor 31P spectra online and stop
exercise upon PCr drop, which may require modifications of the
scanner. Short, intense-exercise protocols have been proposed 2,
as well as exercising a sufficiently large mass of muscle tissue 5
or exercising predominantly oxidative muscles, e.g. tibialias
anterior 6. Here, we explored the possibility to lower the
exercise intensity (force and frequency) and duration, to analyse the
outcome with low-exercise protocols on the frequently studied – but
predominantly glycolytic – plantarflexor muscle gastrocnemius
medialis (GM), and to test what depletion is necessary to quantify
τPCr.Methods
The metabolic and haemodynamic exercise response was simultaneously
assessed in the calf, along GM, using multi-nuclear (1H/31P) data
acquisition with high spatio-temporal resolution. Localised
multi-voxel semi-LASER 31P MRS (VOIs = 25 cm3,
TE = 29 ms) was
interleaved
with multi-slice
1H arterial spin labelling MRI (FAIR ASL,
TE = 20 ms, 10
slices, d = 6 mm),
yielding a multi-nuclear
dataset every TR = 6 s. The
protocol 7
was applied using a custom-built 31P/1H coil array in a 7 T MR Scanner
(Siemens, Erlangen, Germany).
Five young healthy
subjects (4 male / 1 female) exercised twice in one scan session,
each. Exercise force was of 15 – 28 % MVC, the prescribed standard
exercise was 2 min, with two pushes per TR on a non-magnetic
ergometer with adjustable force every 6 s; deviations from standard
protocol see results.Results
With this low-intensity exercise protocol, the 31P spectra localised to GM still showed PCr
depletions of 46 ± 24 % (mean ± SD; range: 15 % – 83 %), very consistently at both voxel
positions, see Fig. 1a. It was possible to reliably quantify τPCr
with depletions as low as 19 %, see Fig. 2. Simultaneously acquired
1H MRI (Fig. 1b)
showed increased perfusion in GM and gastrocnemius lateralis (GL), see Fig. 3,
but this activity was smaller in all evaluated muscles (GM, GL and
soleus) and across all slices than found in recently previously
published data, that were obtained with the same MR sequence but with 3 min exercise at
30 % MVC 7. Different exercise durations were explored (3
min: pHee = 6.97, 1 min: pHee = 7.08), as well pushing once per TR for 3
min (6 % depletion: τPCr not quantifiable). There
was however no correlation between exercise force or power and PCr
depletion and, pHee or τPCr.Discussion
31P MRS results show the well-known pH decrease above ~60 % depletion
(Fig. 2b) with comparable exercise protocols 3, associated
with an increase of τPCr in acidified tissue (Fig. 2c). Interestingly, very short τPCr was found for
small PCr depletions, particularly when pHee was alkaline due to PCr
splitting. These short τPCrs were reliably
quantified and similar in both voxels, placed adjacently in the same muscle (Fig.
1a, 3a). PCr depletion was matched by an increase in
inorganic phosphate (Pi; data not shown) during exercise,
and Pi decayed post exercise with similar time constants as PCr
recovery, also for the shortest τPCrs. That PCr
depletion and other parameters indicative for activation of GM
were independent from prescribed force indicates that recruitment
patterns are individually very different, especially at such small work levels.
Apparently, exercise load can be distributed to other calf muscles than
GM,
although this muscle bears the main load with
stronger exercise and a straight knee 4,8. Fast 31P
spectroscopic imaging methods may resolve this issue if sufficient
SNR is provided, while unlocalised MRS or large techniques with large
VOIs may rather conceal differences in recruitment, potentially
masking differences in pH and τPCr. To study the fast
PCr recovery found with neutral to alkaline pHee, further studies employing shorter repetition times are desirable.Conclusion
Using low-force exercise protocols, small PCr depletions and neutral
or even alkaline pH can be reached after 2 minutes. Even with
depletions less than 20 %, the PCr recovery time remains reliably
quantifiable and, given no acidification, was found to be as short as
~10 s or less, in several datasets.Acknowledgements
Funded by Austrian Science Fund (FWF) project I 1743-B13 to M.M.References
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