We used the line-scan technique to measure in-vivo diffusion at 7T within human primary somatosensory cortex (S1) and primary motor cortex (M1), achieving voxel sizes as low as 0.25 mm in the radial direction (i.e., orthogonal to the cortical surface). Our results are consistent with recent reports of predominantly tangential diffusion in S1 and, to a lesser extent, radial diffusion in M1; however, the smaller voxel sizes used in our study alleviate concerns regarding partial-volume effects and, perhaps more importantly, enable the study of fine-scale variations in diffusion structure across cortical layers.
Two healthy volunteers (1F/1M, ages: 25–26 years), having given informed consent, were scanned on a Siemens 7T whole-body scanner using a custom-built 31-channel head receive coil and birdcage transmit coil. On each volunteer, a 0.75×0.75×0.75 mm3 FOCI-ME-MPRAGE scan[6] was acquired in ~8 minutes with the following parameters: FA = 7°, TE1/TE2/TI/TR = 1.76/3.7/1100/2530 ms, GRAPPA acceleration factor = 2. A single line was acquired with the line-scan diffusion sequence[3-5], prescribed as perpendicularly as possible to primary somatosensory cortex S1 and primary motor cortex M1 (Fig. 2), using the MPRAGE images as anatomical localizers. The line-scan acquisition parameters were as follows: TR/TE = 2000/50 ms, BW ≈ 120 Hz/pixel, 21 b-values equally spaced from “0” to 1000 s/mm2 (∆/δ = 21.77/16.79 ms and with G = 54.6 mT/m for b = 1000 s/mm2) and three diffusion directions: one parallel to the line, and two perpendicular to the line and to one another. We acquired one run with 0.5 mm voxel size along the line (~2 minute acquisition) and four runs with 0.25 mm voxel size (~8 minutes), with a 256 mm readout field-of-view and nominal line thickness of 3 mm in each case.
To remove shot-to-shot phase variations, the following scheme was used: for each voxel, the average complex-valued signal over a 16 mm window was computed, and the phase of this average was subtracted from the phase of the voxel, on a per-coil basis and for each b-value and each run. The resulting phase-corrected complex signal was then averaged across runs prior to root-sum-of-squares coil combination. The monoexponential model was then fitted per-voxel to the resulting signal versus the 19 b-values between 100 and 1000 s/mm2, using the Levenberg-Marquardt nonlinear least-squares algorithm in MATLAB, yielding an apparent diffusion coefficient (ADC) for each direction. The y-intercept of the fit divided by the standard deviation of the residuals provided an estimate of the SNR at b = 0 for each voxel and diffusion direction.
[1] Hubel DH. Eye, Brain, and Vision. Scientific American Library 1988.
[2] Vogt C, Vogt O. Allgemeinere ergebnisse unserer hirnforschung. J Physiol Neurol (Leipz.) 1919;25:279–462.
[3] Gudbjartsson H, Maier SE, Mulkern RV, Mórocz IA, Patz S, Jolesz FA. Line scan diffusion imaging. Magn Reson Med 1996;36:509–519.
[4] Mulkern RV, Gudbjartsson H, Westin CF, Zengingonul HP, Gartner W, Guttmann CR, Robertson RL, Kyriakos W, Schwartz R, Holtzman D, Jolesz FA, Maier SE. Multi-component apparent diffusion coefficients in human brain. NMR Biomed 1999;12:51–62.
[5] Maier SE, Mulkern RV. Biexponential analysis of diffusion-related signal decay in normal human cortical and deep gray matter. Magn Reson Imaging 2008;26:897–904.
[6] Zaretskaya N, Fischl B, Reuter M, Renvall V, Polimeni JR. Advantages of cortical surface reconstruction using submillimeter 7 T MEMPRAGE. Neuroimage 2017;in press. PMID: 28970143.
[7] Anwander A, Pampel A, Knösche TR. In vivo measurement of cortical anisotropy by diffusion-weighted imaging correlates with cortex type. Proc Intl Soc Mag Reson Med 2010;18:109.
[8] McNab JA, Polimeni JR, Wang R, Augustinack JC, Fujimoto K, Stevens A, Triantafyllou C, Janssens T, Farivar R, Folkerth RD, Vanduffel W, Wald LL. Surface based analysis of diffusion orientation for identifying architectonic domains in the in vivo human cortex. Neuroimage 2013;69:87–100.