Phillip G D Ward1,2,3, Francesco Sforazzini1, Sharna D Jamadar1,2,3, Jakub Baran1, Shenpeng Li1,4, Zhaolin Chen1,4, and Gary F Egan1,2,3
1Monash Biomedical Imaging, Monash University, Melbourne, Australia, 2Monash Institute of Cognitive and Clinical Neurosciences, Monash University, Melbourne, Australia, 3Centre of Excellence for Integrative Brain Function, Australian Research Council, Melbourne, Australia, 4Department of Electrical and Computer Systems Engineering, Monash University, Melbourne, Australia
Synopsis
The
objective was to separate the MRI BOLD response into local and non-local
components using a simultaneously acquired functional FDG-PET (fPET) dataset. We
used a visual task, with both high-frequency and low-frequency temporal components,
to simultaneously evoke glucose and BOLD responses. Joint analysis of the fMRI
and fPET identified two components, including a positively-correlated map of
the visual cortex, and a negatively-correlated map of sub-regions of the visual
cortex in fMRI and the draining vasculature in fPET. These findings provide
preliminary evidence that we can deconstruct the fMRI BOLD signal into local (neuronal)
and non-local (haemodynamic) components using simultaneous fMRI-fPET.
Introduction
The in-vivo
monitoring of neuronal activity at high spatial and temporal resolution is critical
for studies of brain function. The blood-oxygen level dependent (BOLD)
response, measured using functional-MRI (fMRI), is an indirect measure of neuronal
activity derived from the hemodynamic response of the vasculature1. BOLD-fMRI provides excellent spatial (~1-3mm)
and moderate temporal resolution (~1-2seconds); however, it may lack specificity.
BOLD response contributes to the signal in the capillary bed, and in the venous
system as blood drains. As such, there is potential for non-local contamination
in the veins. 18-F Flurodeoxyglucose PET (FDG-PET) provides a direct measure of
neuronal glucose metabolism that is coupled to synaptic activity. FDG-PET is
typically performed statically, with a single injected-bolus of tracer. As
such, it has no effective temporal resolution. In this study, we aim to differentiate
the local and non-local BOLD components using a novel, slow infusion functional
FDG-PET (fPET) acquired simultaneously with BOLD-fMRI data during a visual stimulus
paradigm. Methods
fPET was
acquired simultaneously with fMRI during an embedded block design flickering
(8Hz) checkerboard visual task (Figure 1). The low-frequency on/off design (10-minute
blocks) provided FDG-PET contrast, whilst the high-frequency (32/16-seconds)
design provided BOLD-fMRI contrast. Two periods of full checkerboard, and two
periods of half-checkerboard were performed with periods of rest in between.
Analysis was performed using the first full checkerboard.
All data
was acquired on a 3T Biograph mMR (Siemens, Erlangen) with 90-minutes of PET
list-mode data collection. A continuous infusion of FDG (100MBqs) was
administered at a rate of 36mL/hr. MRI
included: dual-UTE, T1-MPRAGE, T2-FLAIR, GRE, ASL, and fMRI (TR=2450ms, TE=30,
64x64x44, res=3x3x3mm3).
fMRI pre-processing
included B0-unwarping, motion correction and high-pass filtering (0.01 Hz). The
data was de-noised using first level ICA, followed by registration to MNI, and smoothing
(5mm FWHM).
List-mode
data were retrospectively reconstructed into 90 bins (1-minute each), using the
OP-OSEM algorithm with point spread function correction and 5mm Gaussian smoothing. Data was corrected for
attenuation (pseudo-CT μ-maps2), motion3, registered to a PET template, and normalised
to the mean grey-matter activity.
Analysis
was performed at two levels. Firstly, two separate fMRI and fPET group ICA4 (with 7 components) were performed
and the visual cortex activation map identified. Dual regression was
performed (FSL5) to obtain a Z-score map per
subject. A joint ICA was then performed (5 components, Fusion ICA Toolbox6), combining the Z-score maps from
fMRI and fPET into a 4D matrix per subject.
The two main joint components were analysed, with the second component overlaid
upon a vein atlas7 for visual inspection.
Results
Independent
activation maps of the visual cortices were produced for the separate fMRI and
fPET analyses (Figure 2). The joint ICA provided two distinct components, with
the remaining three showing minimal structure. The first joint component showed
a positive correlation between the modalities in the visual cortices, and had a
similar spatial extent as the independent BOLD-fMRI and FDG-fPET activation maps (Figure
3). The second joint component demonstrated a negative correlation between the modalities,
with a predominantly positive BOLD-fMRI signal correlated with a negative FDG-fPET
signal. The spatial extent of the BOLD-fMRI response in the second joint ICA component
was located in the visual cortices but with a smaller spatial extent than identified
in the first joint ICA component. The negative FDG-fPET response was identified
in the venous vasculature (Figure 5). Discussion
Activation
in the visual cortices was mapped using an embedded block design using two
distinct imaging modalities, BOLD-fMRI and FDG-fPET. Joint analysis of the
simultaneous fMRI-fPET demonstrated the expected increased glucose and oxygen in
the activated tissue. This demonstrates the feasibility of slow-infusion FDG
for mapping neuronal responses to slowly varying tasks. We interpret the negatively-correlated
component as a non-tissue venous vasculature signal. Following the neuronal activation,
the draining blood vessels have an increased oxygen content resulting from the
hemodynamic response function to neural activity, and a deceased glucose
content, due to the increased uptake of glucose into brain tissue.Conclusion
In this
work, we have demonstrated a novel task and an imaging strategy to (a) allow
simultaneous high temporal resolution FDG-fPET/BOLD-fMRI imaging, and (b) deconstruct
the BOLD-fMRI signal into neural tissue and vascular components. The simultaneous
acquisition and analysis of the FDG-PET and BOLD-fMRI signals may be useful to
enhance the spatial specificity of the BOLD response, by joint analysis with the
FDG-fPET signal, whilst retaining the high spatial and temporal resolution of
the BOLD-fMRI. In future, we aim to routinely acquire and reconstruct BOLD-fMRI
time-series in which the non-local vascular effects have been removed to provide
purely local neuronal tissue response time-series datasets. Acknowledgements
We would like to thank the volunteers for their time and willingness to participate in this study. We would also like to thank the technical staff, Richard McIntyre and Alexandra Carey, and research assistants, Winnie Orchard and Irene Graafsma, for helping collect the data.References
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