Lanette J Friesen-Waldner1, Conrad P Rockel1, Kevin J Sinclair1, Trevor P Wade1,2, Lauren Smith1, Mohamed Moselhy1, Cheryl Vander Tuin3, Albert P Chen4, Barbra de Vrijer5,6,7, Timothy RH Regnault3,5,6,7, and Charles A McKenzie1,2,6,7
1Medical Biophysics, Western University, London, ON, Canada, 2Robarts Research Institute, London, ON, Canada, 3Physiology and Pharmacology, Western University, London, ON, Canada, 4GE Healthcare, Toronto, ON, Canada, 5Obstetrics and Gynaecology, Western University, London, ON, Canada, 6Children's Health Research Institute, London, ON, Canada, 7Lawson Research Institute, London, ON, Canada
Synopsis
Intrauterine
growth restriction (IUGR) is associated with impaired placental metabolism and
transport. Hyperpolarized carbon-13 (HP13C)
MRI was used to detect metabolic differences between control and IUGR placentae
in pregnant guinea pigs, and to determine the impact of maternal hyperoxygenation
on placental metabolism. Area under the
curve (AUC) was calculated for lactate, alanine, and bicarbonate and expressed
as a ratio relative to pyruvate AUC. The
ratio of alanine to pyruvate AUC decreased significantly in IUGR versus control
placentae. Maternal hyperoxygenation resulted
in significant increases in ratios of alanine and bicarbonate to pyruvate AUCs
for both control and IUGR placentae.
Introduction
Intrauterine growth restriction (IUGR) is
associated with up to a 100-fold increase in perinatal mortality rates1.
It is typically associated with placental insufficiency resulting in impaired
placental gaseous exchange and nutrient transport. Hyperpolarized carbon-13 MRI
(HP13C MRI) can be used to monitor metabolism in placentae2,3,4.
Moreover, maternal hyperoxygenation has been used in IUGR to determine
placental oxygenation response5. We propose that HP13C
MRI can provide a means of assessing metabolic function in IUGR placentae and
that IUGR placentae will respond differently than control placentae to maternal
hyperoxygenation.Objectives
1) To detect
metabolic differences between control and IUGR placentae in pregnant guinea
pigs using hyperpolarized [1-13C]pyruvate MRI, and 2) to determine
the impact of maternal hyperoxygenation on placental metabolism.
Methods
Under a
protocol approved by the institution’s Animal Care Committee, six pregnant
guinea pigs underwent surgery at mid-gestation (31-33 days gestation, term ~69
days). Two sows (5 placentae) had vascular occluders placed on the uterine
artery to restrict uteroplacental blood flow.
The remaining four sows (12 placentae) underwent sham surgery.
MR imaging occurred
at 53-64 days gestation (term 68 days) using a 3T MRI system (Discovery MR750, GE
Healthcare). Animals were anaesthetized
using isoflurane and air, a catheter was placed in the tarsal vein of the
maternal hind paw. T1-weighted 1H images were obtained using a
32 coil cardiac array. HP13C
imaging was performed with a custom-built 13C birdcage
coil (Morris Instruments). [1-13C]pyruvic
acid (Cambridge Isotope Laboratories) containing 15mM OX63 trityl radical
(Oxford Instruments) and 1mM ProHance (Bracco) was hyperpolarized using a DNP
polarizer (Hypersense, Oxford Instruments).
3.5mL of the hyperpolarized solution (80mM [1-13C]pyruvate,
here on referred to as pyruvate) was injected over 12s. Time resolved 3D 13C images were
acquired using a modified IDEAL pulse sequence6 (FOV=20cm,
slice=8.5mm, matrix=24x24x14, BW=+/-8.9kHz, echoes=8, ETL=4, TE1=4.2ms,
ΔTE=1.1ms). A double Gaussian radiofrequency
pulse was used to excite pyruvate with a 1.5° flip angle, and the metabolic
by-products with flip angles of 7.5°. Acquisition started 7.5s after the
beginning of injection and images were acquired every 7.5s for a total of 45s. Maternal
intake was changed from air to 100% oxygen (O2, hyperoxygenation)
and 72+/-5min later the pyruvate injection and imaging were repeated. Sows were then recovered.
Fetal weight <80g (determined at
collection at 65d gestation) was considered IUGR. HP13C images were normalized to
peak pyruvate signal in each acquisition. Pyruvate, lactate, alanine, and
bicarbonate 13C signal intensities were determined in ROIs
encompassing total placental volumes using 3D Slicer (www.slicer.org). Area under the curve (AUC) was calculated for
each metabolite and expressed as a ratio relative to pyruvate7. Unpaired t-tests were performed to determine
significance for control versus IUGR and maternal air versus O2 for
each metabolite using Prism (v7.03, GraphPad).
Results
Eight fetuses in the sham surgery guinea pigs were >80g and two
fetuses in the occluder surgery animals were <80g resulting in eight control
and two IUGR placentae. Pyruvate,
lactate, alanine, and bicarbonate were observed in all placentae (Figures 1 and
2). Significant differences in ratios of
alanine to pyruvate (p<0.001) and bicarbonate to pyruvate (p<0.05) AUCs were
observed between maternal air and hyperoxygenation conditions for both control and
IUGR (Figure 3). A higher ratio of lactate to pyruvate AUCs, and lower ratios
of bicarbonate and alanine to pyruvate AUCs were observed for IUGR compared to
control placentae, but only the ratio of alanine to pyruvate AUC was
significantly different (p<0.001) (Figure 2 and 4).Discussion
Previous reports have identified placental metabolism of pyruvate to
lactate in HP13C MRI2,3,4. This study, to our knowledge,
is the first to identify placental metabolism of pyruvate to lactate, alanine, and
bicarbonate in HP13C images.
Changes in ratios for
lactate, alanine and bicarbonate to pyruvate AUCs with changes in maternal oxygenation
indicate that it is possible to use HP13C MRI to measure a change in
placental metabolism affected by maternal oxygen challenge. These changes were
not significantly different for control compared to IUGR placentae; however,
further experiments are needed to increase the number of IUGR placentae in this
study. The reduction of alanine in
the IUGR placenta suggests a movement of placental pyruvate into either acetyl-CoA
and the TCA cycle or to lactate.Conclusions
HP13C
MRI detected a significant decrease in ratio of alanine to pyruvate AUC in IUGR
guinea pig placentae as compared to control.
By exposing a pregnant guinea pig to hyperoxygenation, significant increases in ratios of alanine and bicarbonate to pyruvate AUCs were observed and
this effect was not different for control compared to IUGR placentaeAcknowledgements
National
Institutes of Health, U01 HD087181-01 and Canadian Institutes of Health
Research, MOP-209113References
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