Mor Mishkovsky1,2, Brian Anderson3, Magnus Karlsson4, Mathilde H Lerche4, A Dean Sherry3, Rolf Gruetter1,5,6, Zoltan Kovacs3, and Arnaud Comment2,7
1Laboratory for Functional and Metabolic Imaging, Ecole Polytechnique Federale de Lausanne (EPFL), Lausanne, Switzerland, 2Institute of Physics of Biological Systems, Ecole Polytechnique Federale de Lausanne (EPFL), Lausanne, Switzerland, 3Advanced Imaging Research Center, University of Texas Southwestern Medical Center, Dallas, TX, United States, 4Albeda Research, ApS, Copenhagen, Denmark, 5Department of Radiology, Universite de Lausanne, Lausanne, Switzerland, 6Department of Radiology, Geneva University Hospital and Faculty of Medicine, Geneva, Switzerland, 7General Electric Healthcare, Buckinghamshire HP8 4SP, United Kingdom
Synopsis
Real-time
glucose metabolism was observed in healthy mice brain following infusion of
hyperpolarized [U-2H, U-13C]glucose and [U-2H,
3,4-13C]glucose. The evolution of lactate formation was readily
observed. In addition, two glycolysis metabolites, namely 3-phosphoglycerate
and pyruvate, were identified. Abnormalities in cerebral glucose metabolism is
associated with large number of diseases so implementation of this method may
prove useful in imaging brain metabolism in various animal models.
Introduction
Glucose
is the primary fuel for the mammalian brain and variations in its metabolism
are often indicative of pathologies
1. Collection of MRS
data during continuous infusion of thermally-polarized
13C-labeled
glucose is a method of choice to study the kinetics of glucose metabolism and
to highlight specific metabolic pathways
2-4.
Hyperpolarization by dissolution DNP offers dramatic enhancement of the
13C
signal in small molecules
5 and offers the
possibility of detecting instantaneous metabolic transformations
6. Despite the
relatively short T
1s of the
13C carbons in the glucose molecule, recent studies
demonstrated the applicability of hyperpolarized [U-
2H, U-
13C]glucose
to monitor metabolism in various
in vitro
systems
7-11.
Hyperpolarized
13C-labeled glucose has also been imaged in rats
in vivo12. Real-time
hyperpolarized [U-
2H, U-
13C]glucose metabolism
in vivo has been reported in lymphoma tumors, but glucose
metabolism could not be directly monitored in the brain
13.
The aim of the present study is to demonstrate that it is indeed possible to
detect real-time cerebral metabolism of hyperpolarized glucose with a time
resolution of sub-seconds.
Methods
[U-2H,
U-13C]glucose and [U-2H, 3,4-13C]glucose
samples were dynamically polarized at 7 T polarizer (196 GHz / 1.00±0.05 K) using
trityl radical as polarizing agent and were dissolved with superheated D2O.
All MR measurements were performed on a 9.4T/31cm actively shielded animal
scanner (Varian/Magnex). The liquid polarization of hyperpolarized [U-2H, U-13C]glucose was measured
inside an injection pump placed at the bore of the MR scanner, by comparing the
hyperpolarized signal to its thermal counterpart (Figure 1). In vivo measurements were performed on C57BL/6J
female mice. Animals were anesthetized using 1.5% isoflurane, a catheter was
placed at their femoral vein and connected to a separator/infusion pump for
automatic infusion of hyperpolarized solutions14. Field
inhomogeneity was corrected using the FASTMAP protocol. In vivo data was
collected using a home-built quadrature-1H single loop 13C
surface coil. Acquisition was triggered
5.5s after bolus injection of hyperpolarized [U-2H, U-13C]glucose
solution (n = 4) and hyperpolarized [U-2H, 3,4-13C]glucose (n = 2) using frequency selective RF pulses of 20°
flip angle in the carboxyl area and 1.4° flip angle in the glucose resonances
every 500 ms. Glucose levels were quantified from blood collected from the tail
vein before and just after i.v. infusion. Animal physiology was measured during
the entire length of the experiments. Lactate-to-pyruvate ratio was calculated
from the area under the curve in the summed spectra. Results
An average 13C
glucose polarization of about 20% was measured in the liquid state at the time
of injection (Figure 1). A typical time course of 13C spectra
detected in the mice brain following hyperpolarized [U-2H, U-13C]glucose
infusion is presented in Figure 2 (lower panel). The formation of [1-13C]lactate was readily
detected at 183.5 ppm with good SNR for quantification (Figure 2 upper panel). Upon
infusing of [U-2H, 3,4-13C]glucose, the lactate doublet
is reduced to a single peak (Figure 3). Two additional metabolites could be
identified in the summed spectra (Figure 3). The resonance at 171.1 ppm corresponds
to the C1 of pyruvate with lactate-to-pyruvate ratio of 17.6 ± 2 (n = 5). The peak at 179.8 ppm was assigned to C1 of 3-phosphoglycerate
(3PG). The blood glucose concentration did not exceed the typical values as in
glucose tolerance test (GTT) with a peak at 16.8±2.8 mM post injection.Discussion
We demonstrate
the feasibility of using hyperpolarized 13C-glucose to measure in
vivo cerebral metabolism in healthy mice using a dose of glucose similar to
that used in a GTT. Replacing [U-2H, U-13C]glucose with
[U-2H, 3,4-13C]glucose improves the sensitivity of the
measurement due to lack of 13C-13C coupling. The glycolytic
intermediate 3PG and glycolytic end product pyruvate were identified in vivo
even though those metabolites cannot be detected using thermally polarized 13C
MRS due to their low concentration. The observed lactate-to-pyruvate ratio is in
agreement with its expected value15,16. Lactate signal was identified
10.5 s after glucose infusion, and the time course of lactate formation could
be quantified. This dynamic data could provide a tool study the kinetics of
cerebral glycolysis with sub-second time resolution, and is complementary to information
obtained by thermally polarized 13C MRS where lactate can be
identified only after 20 min post infusion even when using the an optimized
setup4. Abnormalities in
cerebral glucose metabolism are associated with large number of diseases so
this method may prove useful for imaging brain metabolism in various animal
models.Conclusion
13C glucose can be polarized to high levels that enable its
application to study real-time cerebral metabolism in healthy mice.Acknowledgements
This work was
supported by the Swiss National Science Foundation (grant PP00P2_133562 to
A.C), the Centre d’Imagerie BioMédicale (CIBM) of the UNIL, UNIGE, HUG, CHUV,
EPFL, the Leenards and Jeantet Foundations, and the National Institutes of
Health (P41-EB015908 and R37-HL034557 to ADS).References
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