Synopsis
Conscious regulation of emotions is essential
for sound functioning of an individual, while its disruption leads to several
severe symptoms observed in psychiatric disorders like posttraumatic stress
disorder (PTSD) and mild-traumatic brain injury (mTBI). While the brain regions
activated in emotion regulation have been elucidated in prior works, an
understanding of the underlying network has been elusive. Employing an emotion
regulation task, we discovered the network of emotion regulation in healthy soldiers,
and dysregulation in soldiers with comorbid PTSD/mTBI (N=59). Our work is
significant given that we present, for the first time, the evidence for the
network of emotion regulation/dysregulation.
Introduction
Emotions play an important role in our healthy
functioning. Our ability to shape our emotion experience is known as emotional
regulation [1], involving voluntary
modification of emotions elicited in response to exogenous stimuli. Several
functional MRI activation studies have consistently identified the middle
frontal gyrus (MFG), anterior cingulate and insula to be involved in emotion
regulation [1], but their
limitation lies in the inability to explain the interrelationship between these
regions, i.e. connectivity. The brain network of emotion regulation either in
healthy adults or in psychiatric disorders like posttraumatic stress disorder
(PTSD) and mild-traumatic brain injury (mTBI) has been elusive. Emotion
dysregulation has been regarded as a primary cause for several symptoms
observed in PTSD and mTBI [2]. Using fMRI data
collected during an emotion regulation task, we obtained in a data-driven
fashion, the network of emotion regulation in healthy soldiers and
dysregulation in soldiers with comorbid PTSD and chronic mTBI (or
post-concussion syndrome [PCS]).Methods
Fifty-nine
male U.S. Army soldiers were recruited (comorbid PTSD and PCS [PCS+PTSD]=36, combat
controls=37, matched in age, race and education). FMRI data was acquired in a
Siemens Magnetom Verio 3T scanner (EPI sequence, TR/TE=600/30ms, flip-angle=55o,
multiband-factor=2, voxel-size=3.5×3.5×5mm3). The emotion regulation
task (Fig.1) was similar to Urry et.al. [3]. Participants were
presented images which either elicited a neutral or a negative emotional
response. They were instructed to either “maintain” their emotional response,
or “suppress” it (reduce intensity of negative feelings, requiring emotion
regulation). There were 4 task blocks, with 24-trials in each block (4-trials, 6-repetitions).
The comparison between (i) negative-image, suppress-emotion and (ii)
negative-image, maintain-emotion would inform us on the neural mechanisms
underlying emotion regulation.
Standard
pre-processing was performed in SPM [4]
(realignment, smoothing [8mm-kernel], normalization to MNI-space, quality
control). We first performed activation analysis to identify significantly
activated regions during emotion regulation (see Fig.2 for region selection
procedure). Hemodynamic deconvolution was performed [5] on mean time-series
extracted from identified regions, to minimize the non-neural intra-subject HRF
variability [6]. Given that
voluntary emotion regulation is a top-down process [1], we employed
effective connectivity (EC) modeling using Granger causality (GC) [7] to assess
directional causal relationships between identified regions, similar to recent
works [8, 9, 10]. Subject-wise EC
between all regions were obtained, using which the networks of emotion
regulation in healthy soldiers (‘suppress’ vs ‘maintain’ negative emotion) and
its impairment in PCS+PTSD (control vs PCS+PTSD for ‘suppress’ negative emotion
condition) were obtained (p<0.001, Bonferroni corrected) (Fig.3). We provide
novel evidence for the brain networks of both voluntary emotion regulation, and
its dysregulation in a clinical population.Results and Discussion
In this work, we investigated brain
networks of emotion regulation in healthy soldiers, and dysregulation in
comorbid PCS and PTSD. We identified regions activated during the emotion
regulation task, and defined ROIs around the centroid of each of the nine
identified clusters (Fig.4). Performing EC analysis, we identified the
regulation network as having a top-down architecture with the MFG driving the
rest of the network (insula, medial prefrontal regions, amygdala and lateral
parietal regions) (Figs 5a,5b,5c). During dysregulation this network was
imbalanced with reduction in prefrontal connectivity and elevation of
subcortical and lateral parietal connectivity (Figs 5d,5e,5f).
Our network of emotion regulation fits well
with findings from prior studies [1, 11,
12],
which have identified the pivotal role of MFG in the initiation of emotion
regulation. Although amygdala facilitates emotion generation, and medial
prefrontal regions facilitate subconscious emotion regulation like fear
conditioning [1], the MFG initiates
conscious cognitive emotion regulation [1]. MFG is implicated
in executive functions like cognitive control [13], which are necessary
for regulating emotions. Soldiers with PTSD have been shown to have impaired
cognitive functions associated with the MFG [14], as well as impaired
emotional processing [15]. All our directional
connections are traceable to the MFG, implying that MFG could be the source of
emotion regulation [1].
As for dysregulation, the MFG emerged as the
key source of dysfunction in soldiers with PCS+PTSD. All connections emerging
from MFG exhibited reduced connectivity, whose “ripple-effect” culminated in
disinhibition of amygdala, which might translate into elevated retrieval of
undesirable traumatic memories, causing symptoms like flashbacks, trauma
re-experiencing and hyperarousal. This explanation fits well with behavioral
manifestations of these conditions [2].
In summary, we identified the MFG to be
pivotal during emotion regulation in healthy soldiers and dysregulation in
soldiers with PCS+PTSD. Our results are significant
given that these regions are implicated in prior activation studies [16, 17, 11],
but a precise understanding of the underlying network structure and their
causal relationships had not emerged so far.Acknowledgements
The authors acknowledge financial
support for this work from the U.S. Army Medical Research and Materiel Command
(MRMC) (Grant # 00007218). The views, opinions, and/or findings contained in
this article are those of the authors and should not be interpreted as
representing the official views or policies, either expressed or implied, of
the U.S. Army or the Department of Defense (DoD). The funders had no role in
study design, data collection and analysis, decision to publish, or preparation
of the manuscript. The authors thank the personnel at the TBI clinic and behavioral
health clinic, Fort Benning, GA, USA and the US Army Aeromedical Research
Laboratory, Fort Rucker, AL, USA, and most of all, the soldiers who
participated in the study. The authors thank Julie Rodiek and Wayne Duggan for
facilitating data acquisition.References
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