Stefania Evangelisti1, Claudia Testa2, Laura Ludovica Gramegna1,3, Fabio Pizza3,4, David Neil Manners1, Elena Antelmi3,4, Lia Talozzi1, Claudio Bianchini1, Giuseppe Plazzi3,4, Raffaele Lodi1,3, and Caterina Tonon1,3
1Department of Biomedical and NeuroMotor Sciences, Functional MR Unit, University of Bologna, Bologna, Italy, 2Department of Physics and Astronomy, University of Bologna, Bologna, Italy, 3IRCCS Istituto delle Scienze Neurologiche di Bologna, Bologna, Italy, 4Department of Biomedical and NeuroMotor Sciences, University of Bologna, Bologna, Italy
Synopsis
Narcolepsy type 1 (NT1) is a rare and life-long disease, characterized by central
hypersomnia and cataplexy typically triggered by emotions.
NT1 is linked to a selective loss of hypothalamic hypocretin neurons.
To characterise neurodegeneration, we combined pons 1H-MRS
and whole brain structural analysis in a large and homogenous sample of adult NT1
patients.
1H-MRS showed evidence of pontine neuronal
dysfunction, consistent with its key role in REM sleep regulation. Grey matter loss
was detected in brain regions implicated in the disease pathophysiology, including
frontal-prefrontal cortices, putamen nuclei, thalami, hypothalamus, amygdalae, cerebellum,
and widespread subtle
tissue microstructural alterations were also found.
Introduction
Narcolepsy type 1 (NT1) is a rare central sleep
disorder characterized by excessive daytime sleepiness and cataplexy, i.e. sudden
episode of paralysis of voluntary muscle triggered by strong emotions, sleep paralysis,
hypnagogic hallucinations and disrupted nocturnal sleep1. NT1 is
caused by loss of orexin-producing neurons in the postero-lateral hypothalamus,
which project to widespread brain and brainstem areas2.
Most previous proton MRS studies of NT1 patients have focused on neurodegenerative
changes in hypothalamus3,4,5, while in the pons inconsistent
results regarding metabolites concentrations have been found in limited cohorts6,7.
Similarly, previous structural neuroimaging studies have shown controversial
results consistent with the inhomogeneity of study population and image analysis
methods employed8.
The aim of this study was to test for metabolic
alterations in the pons by 1H-MRS, and to brain grey and white matter
alterations by VBM and TBSS analyses, in a large cohort of adult NT1 cases.Methods
We evaluated 44 NT1 patients, who underwent brain MR
in their diagnostic workup to exclude symptomatic forms due to other
neurological diseases. Acquisitions were performed with a 1.5T GE scanner,
equipped with a head coil. The standardized MR protocol included single voxel 1H-MR
spectroscopy (PRESS) with localization in the pons (TR/TE=1500/40ms; NS=512,
volume of interest mean[SD] 1.78[1.09] ml), volumetric T1-w images (TR/TE/TI=12.5/5.1/600ms,
1mm3 isotropic) and diffusion-weighted MRI (TR/TE=10.000/87.5ms, 25 directions,
b-value=900 mm2s−1, voxel=1.25x1.25x4mm).
Not all the patients completed the whole protocol; groups
of sex- and age-matched healthy controls were included in the analysis (Table 1).
Spectral analyses were performed with LC Model 6.3,
and N-acetyl-aspartate (NAA), choline
(Cho) and myo-Inositol (mI) concentrations were expressed relative to creatine
(Cr).
Voxel-based morphometry (VBM) analyses were performed
on T1-w images using SPM 12.0. After a bicommissural realignment and tissue
segmentation, grey matter images were aligned to an iteratively-created study
specific template (DARTEL), registered to the MNI-152 template and spatially
smoothed (gaussian kernel, FWHM 5mm).
Diffusion-weighted data underwent standard
pre-processing and tensor fitting. Voxelwise analysis of tensor parameters was
then performed using TBSS (Tract-Based Spatial Statistics). Fractional anisotropy (FA) images for all
subjects were non-linearly aligned, a mean FA image was created and thinned to
create a skeleton representing the centres of all tracts common to the group.
Each subject's data was then projected onto this skeleton and the resulting
data fed into voxelwise cross-subject statistics. Similar comparisons were
performed on MD (mean diffusivity), AD (axial diffusivity) and RD (radial
diffusivity) maps.
Group comparisons were performed with univariate
analyses (SPSS®); voxelwise comparisons were non-parametric (permutation method)
with age and sex (and TIV for VBM) as nuisance regressors. Statistical
significance was set to p<0.05 (corrected for multiple comparisons). Results
NAA/Cr and NAA/mI were significantly lower in patients compared to controls (Table 2). Figure 1 shows the voxel localization of pons
spectroscopy and two spectra, acquired from a HC subject and an NT1 patient,
showing the reduced NAA content. VBM analysis revealed grey matter density loss for NT1
patients mainly in precentral and postcentral gyri, frontal gyri, frontal pole,
frontal operculum cortex, frontal orbital cortex, lateral occipital cortex,
insular cortex, bilateral putamen nuclei, bilateral thalami, hypothalamus, amygdala,
hippocampus, temporal poles and cerebellum (Figure 2).
MD and AD values were found significantly higher in
NT1 patients, mainly in corpus callosum, frontal white matter, corticospinal
tract and thalamic area (Figure 3). Differences in FA and RD values between
patients and controls were slightly below the significance threshold.Discussion and conclusions
Pons is considered a central structure in the pathophysiology
of NT1, because of the presence of the reticular regions implicated in REM sleep
regulation
9 and circuits involved in muscle paralysis during cataplexy
1, but consistent alterations
have not as yet been observed using advanced MRI techniques.
This
study demonstrated in NT1 patients a neuro-axonal degeneration in the pons (lower NAA/Cr and NAA/mI)
related to its inability in the inhibition of REM sleep. In comparison to
previous pons MRS studies, our cohort size is larger and homogeneous regarding the
presence of cataplexy.
As for the morphological and microstructural aspects,
we found decreased grey matter in all brain regions involved in the pathophysiology
of the disease (Figures 1,2) in particular in the medial prefrontal cortex that
receives excitatory projection from the hypothalamus, in areas involved in the emotional
circuit (insula and hippocampal) and in the cerebellum
10, and TBSS showed
subtle
widespread microstructural alterations rather than focal loss of integrity in multiple
white matter tracts.
Acknowledgements
No acknowledgement found.References
- Scammell
TE. Narcolepsy. N Engl J Med. 2015 Dec 31;373(27):2654-62.
- Peyron C, Tighe DK, van den Pol AN, de Lecea L,
Heller HC, Sutcliffe JG, Kilduff TS. Neurons containing hypocretin (orexin)
project to multiple neuronal systems. J Neurosci. 1998 Dec 1;18(23):9996-10015.
- Tonon
C, Franceschini C, Testa C, Manners DN, Poli F, Mostacci B, Mignot E, Montagna
P, Barbiroli B, Lodi R, Plazzi G. Distribution of neurochemical abnormalities
in patients with narcolepsy with cataplexy: An in vivo brain proton MR spectroscopy
study. Brain Res Bull. 2009 Sep 28;80(3):147-50.
- Poryazova R, Schnepf B, Werth E, Khatami R,
Dydak U, Meier D, Boesiger P, Bassetti CL. Evidence for metabolic
hypothalamo-amygdala dysfunction in narcolepsy. Sleep. 2009 May;32(5):607-13.
- Lodi R, Tonon C, Vignatelli L, Iotti S, Montagna
P, Barbiroli B, Plazzi G. In vivo evidence of neuronal loss in the hypothalamus
of narcoleptic patients. Neurology. 2004 Oct 26;63(8):1513-5.
- Ellis CM, Simmons A, Lemmens G, Williams SC,
Parkes JD. Proton spectroscopy in the narcoleptic syndrome. Is there evidence
of a brainstem lesion? Neurology. 1998 Feb;50(2 Suppl 1):S23-6.
- Bican
A, Bora I, Algin O, Hakyemez B, Ozkol V, Alper E. Neuroimaging in narcolepsy.
Sleep Med. 2010 Feb;11(2):225-6.
- Wada M, Mimura M, Noda Y, Takasu S, Plitman E,
Honda M, Natsubori A, Ogyu K, Tarumi R, Graff-Guerrero A, Nakajima S.
Neuroimaging correlates of narcolepsy with cataplexy: A systematic review.
Neurosci Res. 2018 Mar 23.
- Plazzi G, Montagna P, Provini F, Bizzi A, Cohen
M, Lugaresi E. Pontine lesions in idiopathic narcolepsy. Neurology. 1996
May;46(5):1250-4.
- Tondelli M, Pizza F, Vaudano AE, Plazzi G,
Meletti S. Cortical and Subcortical Brain Changes in Children and Adolescents
With Narcolepsy Type 1. Sleep. 2017 Dec 13.